Dmanisi skull 5

The Dmanisi skull, also known as Skull 5 or D4500, is one of five skulls discovered in Dmanisi, Georgia and classified as early Homo erectus. Described in a publication in October 2013, it is estimated to be about 1.8 million years old and is the most complete skull of a Pleistocene Homo species, and the first complete adult hominin skull of that degree of antiquity. According to researchers, the discovery "provides the first evidence that early Homo comprised adult individuals with small brains but body mass, stature and limb proportions reaching the lower range limit of modern variation."

The skull has been the cause of a paleontological controversy that is still ongoing as of 2017: many hominin fossils thought to be from different species such as Homo rudolfensis or Homo habilis may not have been separate species at all. Rather, they may have been a single evolving lineage.